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Hamburger and Hamilton Stage 36

HH36 (10 days).
Distal segments of limb proportionately much longer.
Length of third toe, from tip to middle of metatarsal joint = 5.4 0.3 mm.
Tapering primordia of claws just visible on termini of toes and on digit 1 of wing.
Primordium ofthe comb appears as prominent ridge with slightly serrated edge along dorsal mid-line of beak.
'Labial groove' clearly visible at tip of upper jaw, but barely indicated on tip of mandible.
Nostril narrowed to a slit.
Length of beak from anterior angle of nostril to tip of bill = 2.5 mm.
Flight feathers conspicuous; coverts just visible in web of wing.
Feather germs cover tibio-fibular portion of leg.
>= 9-10 rows of feather germs between each upper eyelid and dorsal midline.
Sternal tracts prominent, with 3-4 rows on each side of ventral midline when counted in anterior part of sternum,
merging into many rows around umbilicus.
Nictitating membrane covers anteriormost scleral papillae and approaches cornea.
Lower lid has grown to level of cornea.


  1. TXT - Text Format
    The ABSTRACT ONTOLOGY (NOT Staged/Timed) with all pathways to child nodes materialised, in Plain Text only, as a Tree Report.

  2. OBO - OBO Format
    The ABSTRACT ONTOLOGY (NOT Staged/Timed) in OBO Flat File Format, version 1.2, available for viewing/editing using OBO-Edit.


  1. H. Eyal-Giladi and S. Kochav
    From cleavage to primitive streak formation: A complementary normal table and a new look at the first stages of the development of the chick : I. General morphology
    Developmental Biology, Volume 49, Issue 2, April 1976, Pages 321-337

  2. V. Hamburger and H.L. Hamilton
    A series of normal stages in the development of the chick embryo
    Journal of Morphology, 1951, Volume 88, Pages 49-92 (reprinted in Developmental Dynamics, December 1992, Volume 195, Issue 4, pages 231-72)

  3. R. Bellairs and M. Osmond
    The Atlas of Chick Development
    Academic Press, ISBN 0-12-084790-6, 1998

  4. C.D. Stern and P.W.H. Holland (Ed.s)
    Essential Developmental Biology: A Practical Approach
    Oxford University Press, ISBN 0-19-963423-8, 1993

  5. M.A.J. Selleck and C.D. Stern
    Fate mapping and cell lineage analysis of Hensen's node in the chick embryo
    Development, Volume 112, Pages 615-626, 1991