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Hamburger and Hamilton Stage 32

HH32 (7.5 days).
All digits and 4 toes have lengthened conspicuously.
5th toe has disappeared.
Webs between digits and toes thin, contours concave.
Differences in size of individual digits and toes conspicuous.
Anterior tip of mandible reaches beak.
>= 11 rows of feather germs on dorsal surface at level of legs.
1 row on tail distinct, 2nd row indistinct.
Scapular and flight feather germs barely perceptible or absent.
6-8 scleral papillae in two groups; one dorsal and one ventral.
Circle not yet closed.


  1. TXT - Text Format
    The ABSTRACT ONTOLOGY (NOT Staged/Timed) with all pathways to child nodes materialised, in Plain Text only, as a Tree Report.

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    The ABSTRACT ONTOLOGY (NOT Staged/Timed) in OBO Flat File Format, version 1.2, available for viewing/editing using OBO-Edit.


  1. H. Eyal-Giladi and S. Kochav
    From cleavage to primitive streak formation: A complementary normal table and a new look at the first stages of the development of the chick : I. General morphology
    Developmental Biology, Volume 49, Issue 2, April 1976, Pages 321-337

  2. V. Hamburger and H.L. Hamilton
    A series of normal stages in the development of the chick embryo
    Journal of Morphology, 1951, Volume 88, Pages 49-92 (reprinted in Developmental Dynamics, December 1992, Volume 195, Issue 4, pages 231-72)

  3. R. Bellairs and M. Osmond
    The Atlas of Chick Development
    Academic Press, ISBN 0-12-084790-6, 1998

  4. C.D. Stern and P.W.H. Holland (Ed.s)
    Essential Developmental Biology: A Practical Approach
    Oxford University Press, ISBN 0-19-963423-8, 1993

  5. M.A.J. Selleck and C.D. Stern
    Fate mapping and cell lineage analysis of Hensen's node in the chick embryo
    Development, Volume 112, Pages 615-626, 1991